The rubber gasket on the reservoir? Is this one under the reservoir cap? There were lots of those on Detroit reservoirs, but I don't think they mattered.
The important thing you need to make sure is that the over flow tube from the radiator neck basically forms a continuous, leak-free path into the coolant inside the reservoir. This way when the engine cools, the radiator doesn't suck air back but it sucks the coolant in the reservoir.
This means that the rubber hose connections (to the radiator neck, reservoir) should NOT leak. Use a clamp (worm-gear or spring) carefully if you need so you don't crack the plastic.
If your engine cools down and the air pocket grows then there is a leak somewhere you need to find and fix. Otherwise you'll be working on the water pump and head gasket among the things.
The important thing you need to make sure is that the over flow tube from the radiator neck basically forms a continuous, leak-free path into the coolant inside the reservoir. This way when the engine cools, the radiator doesn't suck air back but it sucks the coolant in the reservoir.
This means that the rubber hose connections (to the radiator neck, reservoir) should NOT leak. Use a clamp (worm-gear or spring) carefully if you need so you don't crack the plastic.
If your engine cools down and the air pocket grows then there is a leak somewhere you need to find and fix. Otherwise you'll be working on the water pump and head gasket among the things.